Flora of New York/Lycophytes

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Class Lycopodiopsida contains the lycophytes (clubmosses, firmosses and quillworts).^[1]^[2]

tracheophytes

lycophytes

Lycopodiales : Lycopodiaceae

Lycopodielloideae

	Lycopodiella spp. (5 bog cubmosses)

	Pseudolycopodiella caroliniana (slender bog clubmoss)

Lycopodioideae

Dendrolycopodium spp. (3 tree cubmosses)

Diphasiastrum spp. (8 ground cedars)

	Spinulum spp. (3 bristly clubmosses)

	Lycopodium spp. (2 running clubmosses)

Huperzioideae

	Huperzia spp. (4 firmosses)

Isoetales : Isoetaceae	Isoetes spp. (8 quillworts)

Selaginellales : Selaginellaceae	Selaginella spp. (3 spikemosses)

euphyllophytes

	Polypodiopsida (ferns)

	spermatophytes (seed plants)

↑ P.F. Stevens, (2001-2015). "Ferns s.l." Angiosperm Phylogeny Website. Page last updated: 03/11/2021
↑ PPG I (2016), A community‐derived classification for extant lycophytes and ferns. Jnl of Sytematics Evolution, 54: 563-603. https://doi.org/10.1111/jse.12229

Class	Order	Family	Subfamily	Genus	Common names	#
Lycopodiopsida (lycophytes)	Lycopodiales	Lycopodiaceae	Lycopodielloideae	Lycopodiella	bog clubmoss	5
			Lycopodielloideae	Pseudolycopodiella	false bog clubmoss	1
			Lycopodioideae	Lycopodium	clubmoss	2
				Dendrolycopodium	tree clubmoss	3
				Spinulum	interrupted clubmoss	3
				Diphasiastrum	ground cedar	7
			Huperzioideae	Huperzia	firmoss	5
	Isoetales	Isoetaceae	—	Isoetes	quillwort	8
	Selaginellales	Selaginellaceae	—	Selaginella	spikemoss	3

Order Lycopodiales

The Lycopodiales ...

Family Lycopodiaceae

The Lycopodiaceae (clubmoss family) can, for identification purposes in New York State, roughly be divided into clumping (firmoss) and creeping (clubmoss) groups.

Lycopodiaceae

Huperzioideae (firmoss)

	Huperzia lucidula (shining firmoss)

	Huperzia appressa (mountain firmoss)

	Huperzia selago (northern firmoss)

	Huperzia × bartleyi (Bartley’s firmoss)

Lycopodielloideae

bog clubmoss

	Lycopodiella inundata (northern bog-clubmoss)

	Lycopodiella appressa (southern bog-clubmoss)

	Lycopodiella alopecuroides (foxtail bog-clubmoss)

slender bog-clubmoss

Pseudolycopodiella caroliniana (Carolina clubmoss)

Lycopodioideae

running clubmoss

	Lycopodium clavatum (staghorn clubmoss)

	Lycopodium lagopus (one-cone clubmoss)

tree clubmoss

	Dendrolycopodium obscurum (flat-branched tree clubmoss)

	Dendrolycopodium dendroideum (prickly tree clubmoss)

	Dendrolycopodium hickeyi (Hickey’s tree clubmoss)

bristly clubmoss

	Spinulum annotinum (common bristly clubmoss)

	Spinulum canadense (northern bristly clubmoss)

groundcedar

	Diphasiastrum digitatum (southern ground cedar)

	Diphasiastrum tristachyum (blue ground cedar)

	Diphasiastrum complanatum (northern ground cedar)

	Diphasiastrum sitchense (Sitka ground cedar)

clumping
(firmoss)

creeping
(clubmoss)

Subfamily Huperzioideae

The Huperzioideae, known as firmosses, have also been treated as a separate family Huperziaceae. Firmosses differ from other Lycopodiaceae in that they grow in clumps, instead of producing creeping, horizontal stems like other family members.^[1] Of the three genera in this group, Huperzia is a temperate and arctic genus and is the only one found naturally in New York. Of the other two genera, Phlegmariurus is native to tropical and subtropical regions, and Phylloglossum is native to Australia and New Zealand.

↑ "Family: Huperziaceae - firmoss family" Go Botany 3.5, Native Plant Trust, 2021

Huperzia

Commonly known as firmoss or fir-clubmoss, Huperzia (hoo-per-zee-uh) was named for Johann Peter Huperz, an early 19th century German botanist who specialized in ferns.

The rare Appalachian firmoss was published as Huperzia appalachiana in 1992, but authorities, including Weakly (2023)^[1] consider it to be indistinct from the previously published Huperzia appressa.

↑ Weakley, A.S., and Southeastern Flora Team. 2023. Flora of the southeastern United States. University of North Carolina Herbarium, North Carolina Botanical Garden, Chapel Hill, U.S.A. Dec 23, 2023.

Lycopodiales — Lycopodiaceae — Huperzioideae — Huperzia
Genus Huperzia Bernh. (1801)	Firmoss	N.Y. Status	Images	Distribution	NY NPT
Huperzia lucidula (Michx.) Trevis. 1803. Lycopodium lucidulum Michx. 1875. Huperzia lucidula (Michx.) Trevis. 1903. Huperzia selago ssp. lucidulum Clute 1937. Lycopodium lucidulum var. verum Wherry 1939. Urostachys lucidulus (Michx.) Herter ex Nessel 1939. Urostachys lucidulus var. reclinatus (Dup) Nessel 1965. Huperzia selago ssp. lucidula (Michx.) Á.Löve & D.Löve 1967. Lycopodium lucidulum var. tryonii Mohlenbr. 1999. Huperzia lucidula var. tryonii (Mohlenbr.) Mohlenbr.	Shining firmoss, Shining clubmoss, Shining fir-clubmoss Lycopode brillant, Huperzie brillant	Native, CoC: 5, Secure, S5, G5 FAC-FACW Perennial, Subshrub		NYFA: 5 iNat	USDA-NN POWO FSUS Go Botany VASCAN ARS-GRIN ITIS FNA FNA Tropicos NatureServe IPN Images Wikispecies
Huperzia appressa (Bach.Pyl. ex Desv.) Á.Löve & D.Löve 1827. Lycopodium selago var. appressum Bach.Pyl. ex Desv. 1902. Lycopodium selago f. appressum (Bach.Pyl. ex Desv.) Gelert 1930. Lycopodium appressum (Bach.Pyl. ex Desv.) Petrov in… 1939. Urostachys selago var. appressus (Bach.Pyl. ex Desv.) Herter ex Nessel in… 1956. Lycopodium selago ssp. appressa (Bach.Pyl. ex Desv.) D.Löve 1961. Huperzia appressa (Bach.Pyl. ex Desv.) Á.Löve & D.Löve in… 1968. Lycopodium selago ssp. appressum (Bach.Pyl. ex Desv.) Hultén 1981. Huperzia selago var. appressa (Bach.Pyl. ex Desv.) Ching 1992. Huperzia appalachiana Beitel & Mickel in…	Appalachian firmoss, Appalachian clubmoss, Appalachian fir-clubmoss, Mountain firmoss, Mountain fir-clubmoss Lycopode des Appalaches	Native, CoC: 9, Rare, S3, G4-G5, NYNHP: 3^[1] FACU Perennial, Subshrub		NYFA: 3 NYFA: Essex, Greene iNat GBIF Essex County	USDA-NN COL WFO POWO FSUS Go Botany VASCAN ITIS FNA FNA NatureServe IPN Images Wikispecies
Huperzia selago (L.) Bernh. ex Schrank & Mart. 1753. Lycopodium selago L. 1779. Lycopodium densum Lam. (nom. superfl.) 1805. Plananthus selago (L.) P.Beauv. 1819. Plananthus fastigiatus Opiz 1821. Lycopodium abietiforme Gray 1829. Huperzia selago (L.) Bernh. ex Schrank & Mart. 1831. Lycopodium selago var. densum Wallr. 1866. Selago vulgaris Schur 1923. Urostachys selago (L.) Herter 1942. Lycopodium selago var. typicum Wherry (not validly publ.) 1955. Mirmau selago H.P.Fuchs 1997. Huperzia acuminata (Sugim.) Satou	Northern fir-moss, Fir clubmoss Lycopode sélagine, Huperzie sélagine	Native, CoC: 10, Endangered, NYNHP: 1^[2]		NYFA: 1 NYFA: 5 counties iNat 2 counties	USDA-NN FSUS Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos NatureServe IPN Images Wikispecies
Huperzia porophila (F.E.Lloyd & Underw.) Holub 1900. Lycopodium porophilum F.E.Lloyd & Underw. 1903. Lycopodium selago var. porophilum Clute 1905. Lycopodium lucidulum var. porophilum (F.E.Lloyd & Underw.) Clute 1908. Lycopodium lucidulum f. porophilum (F.E.Lloyd & Underw.) Clute 1923. Urostachys porophilus (F.E.Lloyd & Underw.) Herter 1939. Urostachys lucidulus var. porophilus (F.E.Lloyd & Underw.) Nessel 1965. Huperzia selago var. porophila (F.E.Lloyd & Underw.) Á.Löve & D.Löve 1985. Huperzia porophila (F.E.Lloyd & Underw.) Holub	Rock clubmoss, Rock firmoss, Lloyd's clubmoss Lycopode porophile	N. America native, N.Y. excluded		NYFA-Excluded NYFA: Excluded iNat Carbon County, PA	USDA-NN POWO FSUS VASCAN FNA Tropicos Images Wikispecies
Huperzia occidentalis (Clute) Kartesz & Gandhi 1903. Lycopodium lucidulum f. occidentale Clute 1903. Lycopodium lucidulum f. occidentalis Clute 1932. Lycopodium lucidulum var. occidentale (Clute) L.R.Wilson 1966. Lycopodium selago f. occidentale (Clute) B.Boivin 1966. Lycopodium selago var. occidentale (Clute) B.Boivin 1967. Lycopodium lucidulum var. tryonii Mohlenbr. 1991. Huperzia occidentalis (Clute) Kartesz & Gandhi 1992. Huperzia occidentalis (Clute) Beitel (nom. illeg.)	Pacific Clubmoss ⓘ Western Clubmoss ⓘ Lycopode de l'Ouest ⓘ	N. America native Northwest, N.Y. excluded Perennial, Herb		NYFA-Excluded NYFA: Excluded iNat GBIF Northwest N.A. only	USDA-NN POWO ITIS FNA Tropicos NatureServe BONAP Images Wikispecies
↑ New York Natural Heritage Program. 2024. Online Conservation Guide for Huperzia appressa. Rare, S3/G5. ↑ New York Natural Heritage Program. 2024. Online Conservation Guide for Huperzia selago. Endangered, S1/G5.

Huperzia hybrids

Lycopodiales — Lycopodiaceae — Huperzioideae — Huperzia
Genus Huperzia Bernh. (1801)	Firmoss	N.Y. Status	Images	Distribution	NY NPT
Huperzia × bartleyi (Cusick) Holub Huperzia lucidula × Huperzia porophila 1987. Lycopodium × bartleyi Cusick 1991. Huperzia × bartleyi (Cusick) Holub 1991. Huperzia × bartleyi (Cusick) Kartesz & Gandhi	Bartley's clubmoss	Native, Unranked		NYFA: U NYFA: Ulster County iNat No NY obsevations	USDA-N0 POWO ITIS Tropicos Images Wikispecies
Huperzia × buttersii (Abbe) Kartesz & Gandhi Huperzia lucidula × Huperzia selago 1953. Lycopodium × buttersii Abbe 1991. Huperzia × buttersii Kartesz & Gandhi	Butters' club-moss, Butters' firmoss Lycopode de Butters	N. America native, N.Y. excluded		NYFA-Excluded iNat	USDA-NN POWO Go Botany VASCAN ITIS FNA Tropicos NatureServe Images Wikispecies
Huperzia × protoporophila A.Haines Huperzia appressa × Huperzia lucidula 2003. Huperzia × protoporophila A.Haines	Hybrid rock firmoss	N. America native, No N.Y. reports		NYFA: D iNat	Go Botany VASCAN Tropicos IPN Images Wikispecies
Huperzia × josephbeitelii A.Haines Huperzia appressa × Huperzia selago 2003. Huperzia × beitelii A.Haines 2003. Huperzia × josephbeitelii A.Haines	Beitel's firmoss, Joseph Beitel's clubmoss	N. America native, No N.Y. reports		NYFA: D iNat	USDA-NN Go Botany VASCAN Tropicos IPN Images Wikispecies

Subfamily Lycopodioideae

Lycopodium

The genus name Lycopodium (lī-kə-ˈpō-dē-əm) was derived from the Greek lukos (wolf) and podion (small foot). It is commonly known as running clubmoss.

Also treated as Lycopodium sect. Lycopodium.

Lycopodiales — Lycopodiaceae — Lycopodioideae — Lycopodium
Genus Lycopodium	Running clubmoss	N.Y. Status	Images	Distribution	NY NPT
Lycopodium clavatum L. 1753. Lycopodium clavatum L. 1805. Lepidotis clavata (L.) P.Beauv. 1805. Lepidotis ciliata P.Beauv. 1805. Lepidotis inflexa P.Beauv. 1810. Lepidotis aristatum Humb. & Bonpl. ex Willd. 1925. Lycopodium clavatum var. laurentianum Vict. 1925. Lycopodium clavatum var. subremotum Vict.	Common clubmoss, Running clubmoss, Running pine, Running cedar, Staghorn clubmoss, Elk moss Lycopode à massue	Native, CoC: 5, Secure		NYFA: 5 iNat	USDA-NN POWO Go Botany ARS-GRIN ITIS FNA FNA Images Wikispecies
Lycopodium lagopus (Laest. ex Hartm.) Zinserl. ex Kuzen 1831. Lycopodium clavatum var. monostachyon Hook. & Grev. 1838. Lycopodium clavatum var. integrifolium (Hook. ex Goldie) Spring 1842. Lycopodium clavatum var. integerrimum Spring 1858. Lycopodium clavatum var. lagopus Laest. ex C.Hartm. 1901. Lycopodium clavatum var. brevispicatum Peck 1910. Lycopodium clavatum var. megastachyon Fernald & Bissell 1925. Lycopodium clavatum f. furcatum Vict. 1950. Lycopodium clavatum ssp. monostachyon (Hook. & Grev.) Selander 1953. Lycopodium lagopus (Laest. ex C.Hartm.) Zinserl. ex Kuzen 1958. Lycopodium clavatum ssp. megastachyon (Fernald & Bissell) Á.Löve & D.Löve	One-cone clubmoss, One-cone ground-pine, Running pine, Running cedar, Arctic stag's-horn clubmoss Lycopode patte-de-lapin	Native, CoC: 4, Likely secure		NYFA: 4 iNat	USDA-NN POWO Go Botany ITIS FNA FNA Images Wikispecies

Dendrolycopodium

From Greek δενδρος (denros: tree), λυκος (lukos: wolf), and ποδιων (podion: legs), Dendrolycopodium (den-drō-lī-kə-ˈpō-dē-əm) contains tree-clubmoss species, which are fairly erect and have branched stems. This genus, also treated as Lycopodium sect. Obscura, is characterized by stalked terminal strobili, which contain their sporangia.

Lycopodiales — Lycopodiaceae — Lycopodioideae — Dendrolycopodium
Genus Dendrolycopodium	Tree clubmoss	N.Y. Status	Images	Distribution	NY NPT
Dendrolycopodium obscurum (L.) A.Haines 1753. Lycopodium obscurum L. 1833. Lycopodium dendroideum var. obscurum (L.) Torr. ex Beck 1937. Lycopodium obscurum var. genuinum Wherry 2003. Dendrolycopodium obscurum (L.) A.Haines	Ground pine, Princess-pine, Eastern tree clubmoss, Rare clubmoss, Flat-branch tree clubmoss Lycopode obscur	Native, CoC: 5, Secure FACU		NYFA: 5 iNat	USDA-NN Go Botany ARS-GRIN ITIS FNA FNA Images Wikispecies
Dendrolycopodium dendroideum (Michx.) A.Haines 1803. Lycopodium dendroideum Michx. 1890. Lycopodium obscurum var. dendroideum D.C.Eaton 1916. Lycopodium obscurum var. hybridum Farw. 2003. Dendrolycopodium dendroideum (Michx.) A. Haines	Prickly tree clubmoss, Northern tree clubmoss, Treelike clubmoss, Prickle tree club-moss Lycopode dendroïde	Native, CoC: 5, Secure FACU		NYFA: 5 iNat	USDA-NN Go Botany ARS-GRIN ITIS FNA FNA Images Wikispecies
Dendrolycopodium hickeyi (W.H.Wagner, Beitel, & Moran) A.Haines 1977. Lycopodium obscurum var. isophyllum Hickey 1989. Lycopodium hickeyi W.H.Wagner, Beitel & R.C.Moran 2003. Dendrolycopodium hickeyi (W.H.Wagner, Beitel, & Moran) A.Haines	Pennsylvania clubmoss, Hickey's tree club-moss	Native, CoC: 5, Secure Perennial, Herb		NYFA: 5 iNat	USDA-NN Go Botany ITIS FNA FNA Images Wikispecies

Spinulum

Spinulum (ˈspīn-yə-ləm) is also treated as Lycopodium sect. Annotina.

Lycopodiales — Lycopodiaceae — Lycopodioideae — Spinulum
Genus Spinulum	Bristly clubmoss	N.Y. Status	Images	Distribution	NY NPT
Spinulum annotinum (L.) A.Haines 1753. Lycopodium annotinum L. 1772. Lycopodium dubium Zoëga 2003. Spinulum annotinum (L.) A.Haines	Common bristly clubmoss, Common stiff clubmoss, Interrupted clubmoss Lycopode interrompu	Native, CoC: 5, Secure		NYFA: 5 iNat	USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Images Wikispecies
Spinulum canadense (Nessel) A.Haines 1772. Lycopodium dubium auct. non Zoëga 1834. Lycopodium pungens (Desv. ex Spring) Bach.Pyl. ex Iljin 1842. Lycopodium annotinum var. pungens Desv. 1842. Lycopodium pungens La Pylaie ex Desv. 1843. Lycopodium annotinum var. montanum Tuck. 1939. Lycopodium canadense Romhild ex Nessel 1968. Lycopodium annotinum ssp. pungens Hultén 1969. Lycopodium annotinum ssp. dubium (Zoëga) Kallio, Laine & Y.Mäkinen 2003. Spinulum canadense (Romhild ex Nessel) A.Haines	Northern bristly clubmoss, Northern stiff clubmoss, Northern interrupted clubmoss	Native, CoC: 10, Endangered		NYFA: 1-2 iNat	Go Botany VASCAN Tropicos Images Wikispecies
Spinulum annotinum × canadense Spinulum annotinum × Spinulum canadense	Hybrid bristly clubmoss	Native, Unranked		NYFA: U	Images Wikispecies

Diphasiastrum

Ground cedar (genus Diphasiastrum) is also treated as Lycopodium sect. Complanata.

Lycopodiales — Lycopodiaceae — Lycopodioideae — Diphasiastrum
Genus Diphasiastrum Holub (1975)	Ground-cedar	N.Y. Status	Images	Distribution	NY NPT
Diphasiastrum digitatum (Dill. ex A.Braun) Holub 1848. Lycopodium digitatum Dill. ex A.Braun 1901. Lycopodium complanatum var. flabelliforme Fernald 1911. Lycopodium flabelliforme (Fernald) Blanch. 1965. Diphasium complanatum ssp. flabelliforme in… 1975. Diphasiastrum digitatum (Dill. ex A.Braun) Holub	Southern groundcedar, Southern ground-cedar, Fan club-moss, Southern running-pine Lycopode en éventail, Courants verts, Lycopode aplati variété en éventail	Native, CoC: 4, Secure, S5, G5		NYFA: 5 iNat	USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA FNA Tropicos IPN Images Wikispecies
Diphasiastrum tristachyum (Pursh) Holub 1814. Lycopodium tristachyum Pursh in… 1840. Lycopodium clavatum var. tristachyum (Pursh) Hook. in… 1848. Lycopodium complanatum ssp. tristachyum (Pursh) Dostál in… 1859. Lycopodium complanatum ssp. chamaecyparissus in… 1905. Lycopodium complanatum var. tristachyon Clute in… 1944. Diphasium tristachyum (Pursh) Rothm. in… 1961. Diphasium chamaecyparissus in… 1975. Diphasiastrum tristachyum (Pursh) Holub in…	Blue ground-cedar, Deep-root clubmoss	Native, CoC: 4, Secure, S5, G5		NYFA: 5 iNat	USDA-NN POWO Go Botany ITIS FNA FNA Tropicos Images Wikispecies
Diphasiastrum complanatum (L.) Holub 1753. Lycopodium complanatum L. in… 1805. Lepidotis complanata (L.) P.Beauv. in… 1840. Lycopodium anceps Wallr. in… 1944. Diphasium complanatum (L.) Rothm. in… 1958. Diphasium anceps (Asch.) Á.Löve & D.Löve in… 1975. Diphasiastrum complanatum (L.) Holub in…	Northern groundcedar, Trailing clubmoss, Northern running-pine, Lycopode aplati	Native, CoC: 8, Endangered, S2, G5		NYFA: 1 iNat	USDA-NN Go Botany ARS-GRIN ITIS FNA FNA Images Wikispecies
Diphasiastrum sitchense (Rupr.) Holub (1975) 1845. Lycopodium sitchense Rupr. in… 1916. Lycopodium complanatum var. sitchense in… 1923. Lycopodium × sabinifolium var. sitchense in… 1965. Lycopodium × sabinifolium ssp. sitchense in… 1965. Diphasium sitchense (Rupr.) Á.Löve & D.Löve in… 1975. Diphasiastrum sitchense (Rupr.) Holub in…	Sitka groundcedar, Alaskan clubmoss, Sitka club-moss, Lycopode de Sitka	Native, CoC: 10, Endangered, S1, G5		NYFA: 1 iNat	USDA-NN Go Botany ARS-GRIN ITIS FNA Images Wikispecies
Diphasiastrum × sabinifolium (Willd.) Holub (1975) Diphasiastrum sitchense × Diphasiastrum tristachyum Lycopodium × sabinifolium Willd. Lycopodium sabinifolium Willd. Lycopodium armatum Desv.	Ground-fir, Savinleaf groundpine, Savin-leaved groundpine, Cypress clubmoss	Native, Unranked, SNA, GNA		NYFA: 3 iNat	USDA-NN ARS-GRIN ITIS FNA Images Wikispecies
Diphasiastrum × zeilleri (Rouy) Holub (1975) Diphasiastrum complanatum × Diphasiastrum tristachyum Lycopodium × zeilleri Diphasium zeilleri (Rouy) Damboldt	Zeiller's running-pine, Zeiller's clubmoss	Native, Unranked, SNA, GNA		NYFA: 2 iNat	USDA-NN ITIS FNA Images Wikispecies
Diphasiastrum × habereri (House) Holub (1975) Diphasiastrum digitatum × Diphasiastrum tristachyum Lycopodium × habereri House Lycopodium complanatum var. habereri Lycopodium tristachyum var. habereri	Haberer's clubmoss	Native, Unranked, SNA, GNA		NYFA: U iNat	USDA-NN ITIS FNA Images Wikispecies

Subfamily Lycopodielloideae

Lycopodiella

Bog clubmoss. (Lycopodiella in Latin means little wolf's-foot.)

Lycopodiales — Lycopodiaceae — Lycopodielloideae — Lycopodiella
Genus Lycopodiella Holub (1964)	Bog clubmoss	N.Y. Status	Images	Distribution	NY NPT
Lycopodiella inundata (L.) Holub 1753. Lycopodium inundatum L. 1805. Plananthus inundatus (L.) P.Beauv. 1852. Lepidotis inundata (L.) Opiz 1912. Lepidotis inundata (L.) Börner 1964. Lycopodiella inundata (L.) Holub	Northern bog clubmoss, Marsh clubmoss, Inundated clubmoss Lycopodielle inondé	Native, CoC: 8, Secure OBL Perennial, Subshrub		NYFA: 5	USDA-NN Go Botany ARS-GRIN ITIS FNA Tropicos Images Wikispecies
Lycopodiella appressa (Chapm.) Cranfill 1843. Lycopodium inundatum var. bigelovii Tuck. 1878. Lycopodium inundatum var. appressum Chapm. 1900. Lycopodium appressum F.E.Lloyd & Underw. 1981. Lycopodiella appressa (Chapm.) Cranfill 1985. Lycopodiella bigelowii (Tuck.) Holub	Southern bog clubmoss, Appressed bog-clubmoss, Chapman's club-moss	Native, CoC: 7, Likely secure FACW Perennial, Subshrub		NYFA: 4	USDA-NN Go Botany ITIS FNA Images Wikispecies
Lycopodiella alopecuroides (L.) Cranfill 1753. Lycopodium alopecuroides L. 1944. Lepidotis alopecuroides (L.) Rothm. 1981. Lycopodiella alopecuroides (L.) Cranfill	Foxtail bog-clubmoss, Foxtail clubmoss	Native, CoC: 7, Vulnerable, S2?, G2 FACW Perennial, Subshrub		Kings, Nassau, Richmond, Suffolk NYFA: 3	USDA-N0 Go Botany ITIS FNA Images Wikispecies
Lycopodiella × copelandii (Eiger) Cranfill (1981) L. alopecuroides × L. appressa	Copeland's bog clubmoss, Copeland's clubmoss	Native, Unranked		Suffolk NYFA: U	USDA-N0 ITIS Images Wikispecies
Lycopodiella × gilmanii A.Haines (2003) L. appressa × L. inundata	Gilman's bog clubmoss, Gilman's clubmoss	Native, Unranked		NYFA: U	ITIS Images Wikispecies
Genus Lycopodiella (excluded taxa)	Bog clubmoss	N.Y. Status	Images	Distribution	NY NPT
Lycopodiella × robusta ( R.J.Eaton) A.Haines (2002) L. alopecuroides × L. inundata Lycopodium inundatum var. robustum	Bog clubmoss	Native, N.Y. excluded		NYFA: U	Images Wikispecies

Pseudolycopodiella

Lycopodiales — Lycopodiaceae — Lycopodielloideae — Pseudolycopodiella
Genus Pseudolycopodiella (L.) Holub	False bog-clubmoss	N.Y. Status	Images	Distribution	NY NPT
Pseudolycopodiella caroliniana (L.) Holub 1753. Lycopodium carolinianum L. 1968. Lycopodiella caroliniana (L.) Pic.Serm. 1983. Pseudolycopodiella caroliniana Holub 1983. Pseudolycopodiella meridionalis Holub	Carolina clubmoss, Slender clubmoss, Slender bog club-moss	Native, CoC: 9, Endangered, NYNHP: 1^[1]		NYFA: 1	USDA-N0 Go Botany ARS-GRIN ITIS FNA Tropicos Images Wikispecies
↑ New York Natural Heritage Program. 2024. Online Conservation Guide for Pseudolycopodiella caroliniana. Endangered, S1/G5T4.

Order Selaginellales

Family Selaginellaceae

Selaginella

Three spikemoss (Selaginella) species have been found growing wild in New York.

Selaginellales — Selaginellaceae — Selaginella
Genus Selaginella	Spikemoss	N.Y. Status	Images	Distribution	NY NPT
Selaginella rupestris (L.) Spring 1753. Lycopodium rupestre L. in… 1803. Stachygynandrum rupestre (L.) P.Beauv. in… 1838. Selaginella rupestris (L.) Spring in… 1849. Selaginella rupestris var. borealis Spring in… 1891. Lycopodioides rupestris (L.) Kuntze in… 1900. Selaginella bourgeaui Hieron. in… 1908. Selaginella rupestris f. borealis (Spring) Clute ... 1992. Bryodesma rupestre (L.) Soják in…	Rock spikemoss, Dwarf spikemoss, Northern selaginella Sélaginelle des rochers, Sélaginelle rupestre	Native, CoC: 9, Likely secure, S4 UPL Perennial, Herb		NYFA-4 iNat	NRCS-NN POWO Go Botany ITIS FNA Images Wikispecies
Selaginella apoda (L.) Spring 1753. Lycopodium apodum L. 1840. Selaginella apus Spring 1840. Selaginella apoda (L.) Spring	Meadow spikemoss, Creeping spikemoss Sélaginelle apode	Native, CoC: 6, Unranked, Under review^[1] FACW		NYFA-2? iNat	NRCS-N0 Go Botany ITIS FNA Images Wikispecies
Selaginella eclipes W.R.Buck 1977. Selaginella eclipes W.R.Buck 1996. Selaginella apoda ssp. eclipes Skoda	Hidden spikemoss, Buck's meadow spikemoss, Creeping spikemoss	Native, CoC: 8, Endangered FACW		NYFA-1	NRCS-NN Go Botany ITIS FNA Tropicos Images Wikispecies
Genus Selaginella (excluded taxa)	Spikemoss	N.Y. Status	Images	Distribution	NY NPT
Selaginella selaginoides (L.) P. Beauv. ex Mart. & Schrank 1753. Lycopodium selaginoides L. 1805. S. spinosa P.Beauv. illeg. 1841. S. selaginoides (L.) P.Beauv. ex Mart. & Schrank	Club spikemoss ⓘ Northern spikemoss ⓘ Northern spike-moss ⓘ Prickly mountain-moss ⓘ Lesser-club-moss ⓘ	N. America native, N.Y. excluded FACW Perennial, Herb		NYFA-Excluded	USDA-NN Go Botany ARS-GRIN Images Wikispecies
↑ 2019 Rare Plant Status Lists – Review List - New York Natural Heritage Program

Order Isoetales

Family Isoetaceae

Isoetes

North American Isoetes lacustris plants have recently been segregated as Isoetes macrospora but can't be easily differentiated except by geography.^[1]

↑ W. Carl Taylor, Neil T. Luebke, Donald M. Britton, R. James Hickey, Daniel F. Brunton (2020). FNA Volume 2. Flora of North America Association

Isoetales — Isoetaceae — Isoetes
Genus Isoetes L.	Quillwort	N.Y. Status	Images	Distribution	NY NPT
Isoetes engelmannii A.Braun 1846. Isoetes engelmannii A.Braun in… 1846. Isoetes lacustris var. microspora Engelm. in… 1867. Isoetes engelmannii var. gracilis Engelm. in… 1891. Calamaria engelmannii (A.Braun) Kuntze in… 1905. Isoetes engelmannii var. fontana A.A.Eaton in… 1994. Isoetes × fairbrothersii J.D.Montgom. & W.C.Taylor in…	Engelmann's quillwort, Appalachian quillwort Isoète d'Engelmann	Native, CoC: 6, Secure, S5 OBL Perennial, Herb, Subshrub, diploid (2n = 22)		NYFA: 5 NYFA: 23 counties iNat GBIF Cortland (2019)	USDA-NN COL WFO POWO FSUS Go Botany VASCAN ITIS FNA FNA Tropicos NatureServe BONAP IPN Images Wikispecies
Isoetes echinospora Durieu ssp. muricata A.Löve & D.Löve^[1] 1789. Isoetes setacea Lam. p.p. 1827. Isoetes tenella Léman ex Desv. POWO in… 1864. Isoetes braunii Durieu in… 1867. Isoetes echinospora var. braunii Engelm. 1945. Isoetes muricata var. braunii (Engelm.) C.F.Reed in… 1961. Isoetes echinospora ssp. muricata (Durieu) Á.Löve & D.Löve NYFA	Spiny-spore quillwort, Spring quillwort, Braun's quillwort, Bristlelike quillwort Isoète à spores épineuses, Isoète à spores spinuleuses	Native, CoC: 7, Likely secure, S4 OBL Perennial, Herb, diploid (2n = 22)		NYFA: 4	USDA-? Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos Images Wikispecies
Isoetes tuckermanii A.Braun ex Engelm. 1867. I. tuckermanii A.Braun ex Engelm. 1900. I. tuckermanii var. borealis A.A.Eaton 1981. I. acadiensis Kott 1991. I. tuckermanii ssp. acadiensis Taylor & Luebke	Tuckerman's quillwort Isoète de Tuckerman	Native, CoC: 10, Likely secure, tetraploid (2n = 44)		NYFA: 4	USDA-NN FSUS Go Botany ITIS FNA Images Wikispecies
Isoetes × eatonii R.Dodge Isoetes echinospora × Isoetes engelmannii 1896. Isoetes eatonii R.Dodge 1900. Isoetes gravesii A.A.Eaton Isoetes eatonii var. gravesii A.A.Eaton	Eaton's quillwort	Native, Vulnerable		NYFA: 3	USDA-NN ITIS Images Wikispecies
Isoetes macrospora Durieu 1753. Isoetes lacustris L. in… 1864. Isoetes macrospora Durieu in… 1900. Isoetes hieroglyphica A.A.Eaton 1922. Isoetes macrospora f. hieroglyphica N.Pfeiff. ...	Big-spore quillwort ⓘ Lake quillwort Carved quillwort Warty Quillwort Isoète lacustre Isoète hiéroglyphique	Native, CoC: 10, Rare, S3 OBL Perennial, Herb, decaploid (2n = 110)		NYFA: 3 NYFA: 5 counties iNat GBIF St. Lawrence only	USDA-NN Go Botany ITIS FNA FNA Images Wikispecies
Isoetes septentrionalis D.F.Brunt. 1846. Isoetes riparia var. canadensis Engelm. in… 1903. Isoetes canadensis var. robbinsii A.A. Eaton 2015. Isoetes septentrionalis D.F. Brunt. in…	Northern shore quillwort, Northern quillwort isoète du Nord	Native, CoC: 10, Endangered OBL Perennial, Herb, tetraploid (2n = 44)		NYFA: 1	USDA-N0 FSUS Go Botany VASCAN ITIS FNA Images Wikispecies
Isoetes × robusta (Engelm.) D.F.Brunt. Isoetes engelmannii × Isoetes septentrionalis 1898. Isoetes × dodgei A.A.Eaton auct. Isoetes × brittonii auct. Isoetes engelmannii × riparia	Hybrid of Engelmann's quillwort & northern shore quillwort	Native, No recent reports		NYFA: Z?	USDA-N0 Go Botany ARS-GRIN ITIS Images Wikispecies
↑ The New York Flora Atlas and New England Wild Flower Society list the spiny-spore quillwort as Isoëtes echinospora ssp. muricata because the leaves of the North American plants of bear stomata, while those of the European plants do not. The other sources used here treat ssp. muricata as a synonym.

Isoetes (excluded)

Isoetales — Isoetaceae — Isoetes
Genus Isoetes (excluded taxa)	Quillwort	N.Y. Status	Images	Distribution	NY NPT
Isoetes riparia Engelm. ex A.Braun^[1] 1846. Isoetes riparia Engelm. ex A.Braun 18??. Isoetes saccharata Engelm. in A.Gray 1891. Calamaria saccharata (Engelm.) Kuntze 1901. Isoetes canadensis (Engelm.) A.A.Eaton 1949. Isoetes riparia f. saccharata ( Engelm. ex A.Gray ) Proctor	Shore quillwort, Canada shore quillwort	Native, N.Y. excluded		NYFA-Excluded	USDA-NN Go Botany ITIS Tropicos BONAP Images Wikispecies
Isoetes × harveyi A.A.Eaton Isoetes lacustris × Isoetes tuckermanii 1900. Isoetes × heterospora A.A.Eaton Isoetes tuckermanii var. harveyi Clute	Harvey's quillwort	Native, N.Y. excluded		NYFA-Excluded	USDA-NN Go Botany ITIS FNA Images Wikispecies
Isoetes × novae-angliae D.F. Brunt. & D.M. Britton Isoetes septentrionalis × Isoetes tuckermanii	New England quillwort	Native, N.Y. excluded		NYFA-Excluded	USDA-N0 Go Botany ITIS FNA Images Wikispecies
↑ Isoetes riparia was apparently misapplied to reports of Isoetes septentrionalis in New York. (NYFA (2016)

[1] P.F. Stevens, (2001-2015). "Ferns s.l." Angiosperm Phylogeny Website. Page last updated: 03/11/2021

[2] PPG I (2016), A community‐derived classification for extant lycophytes and ferns. Jnl of Sytematics Evolution, 54: 563-603. https://doi.org/10.1111/jse.12229

[3] "Family: Huperziaceae - firmoss family" Go Botany 3.5, Native Plant Trust, 2021

[4] Weakley, A.S., and Southeastern Flora Team. 2023. Flora of the southeastern United States. University of North Carolina Herbarium, North Carolina Botanical Garden, Chapel Hill, U.S.A. Dec 23, 2023.

[5] New York Natural Heritage Program. 2024. Online Conservation Guide for Huperzia appressa. Rare, S3/G5.

[6] New York Natural Heritage Program. 2024. Online Conservation Guide for Huperzia selago. Endangered, S1/G5.

[7] New York Natural Heritage Program. 2024. Online Conservation Guide for Pseudolycopodiella caroliniana. Endangered, S1/G5T4.

[8] 2019 Rare Plant Status Lists – Review List - New York Natural Heritage Program

[9] W. Carl Taylor, Neil T. Luebke, Donald M. Britton, R. James Hickey, Daniel F. Brunton (2020). FNA Volume 2. Flora of North America Association

[10] The New York Flora Atlas and New England Wild Flower Society list the spiny-spore quillwort as Isoëtes echinospora ssp. muricata because the leaves of the North American plants of bear stomata, while those of the European plants do not. The other sources used here treat ssp. muricata as a synonym.

[11] Isoetes riparia was apparently misapplied to reports of Isoetes septentrionalis in New York. (NYFA (2016)

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